CBS-KNAW fungal biodiversity centre
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Centraalbureau voor Schimmelcultures

Utrecht, The Netherlands

Mycotoxin production by common filamentous fungi

J. C. FRISVAD and U. THRANE - BioCentrum-DTU
Technical University of Denmark, DK-2800 Lyngby, Denmark

For a more detailed description see: Introduction to food borne fungi.

Toxic metabolite production by species in Ascomycotina.
Mycotoxins sensu stricto are marked with an asterix *.
Many other secondary metabolites are produced by most species but not listed here.


1. Eurotiales
2. Sphaeriales
3. Dothidiales
4. Unknown

1. Eurotiales

  • Byssochlamys (anamorph Paecilomyces)
  • B. fulva: patulin*, byssochlamic acid, byssotoxin**
  • B. nivea: patulin*, byssochlamic acid, malformins
  • Pae. variotii I: patulin*
  • Pae. variotii II: viriditoxin*
  • Talaromyces (anamorph Penicillium subgenus Biverticillium)
  • T. macrosporum: duclauxin*
  • P. crateriforme: rubratoxin*, rugulovasine A & B*, spiculisporic acid*
  • P. funiculosum: ?
  • P. islandicum: rugulosin*, luteoskyrin*, islanditoxin*, cyclochlorotine*, erythroskyrin*, emodin*
  • P. rugulosum: rugulosin*
  • P. variabile: rugulosin*
  • P. purpurogenum: ?
  • Eupenicillium (anamorph Penicillium subgenus Aspergilloides, Furcatum and Penicillium)
  • P. atramentoseum: oxaline, rugulovasine A+B
  • P. aurantiogriseum: penicillic acid*, verrucosidin*, nephrotoxic glycopeptides*, anacine, auranthine, aurantiamine
  • P. aurantiocandidum: penicillic acid*, puberulic acid, cyclopenin, cyclopenol
  • P. brasilianum: penicillic acid*, verrucologen*, fumitremorgin A* & B*, viridicatumtoxin*
  • P. brevicompactum: botryodiploidin*, mycophenolic acid, Raistrick phenols, brevianamide A
  • P. camemberti: cyclopiazonic acid*
  • P. carneum: mycophenolic acid, patulin*, penitrem A*, roquefortine C*
  • P. chrysogenum: roquefortine C*, meleagrin, penicillin***, chrysogine
  • P. citrinum: citrinin*, tanzawaic acid A
  • P. commune: cyclopiazonic acid*, rugulovasine A & B, cyclopaldic acid
  • P. corylophilum: ?
  • P. crustosum: roquefortine C*, penitrem A*, terrestric acid, cyclopenin, cyclopenol
  • P. cyclopium: penicillic acid*, xanthomegnin*, viomellein*
  • P. digitatum: tryptoquivalins
  • P. discolor: chaetoglobosin A, B & C*, cyclopenin, cyclopeol
  • P. echinulatum: territrems*, cyclopenin, cyclopenol
  • P. expansum: roquefortine C*, patulin*, citrinin*, communesins, chaetoglobosin C*
  • P. freii: xanthomegnin*, viomellein*, vioxanthin*, penicillic acid*, aurantiamine, cyclopenon, cyclopenol
  • P. glabrum: citromycetin*
  • P. griseofulvum: roquefortine C*, cyclopiazonic acid*, patulin*, griseofulvin*
  • P. hirsutum: roquefortine C*, terrestric acid
  • P. hordei: roquefortine C*, terrestric acid
  • P. italicum: italicic acid, verrucolone
  • P. melanoconidium: penicillic acid*, penitrem A*, roquefortine C*, verrucosidin*
  • P. nalgiovense: penicillin***, chrysogine
  • P. nordicum: ochratoxin A*, anacine, verrucolone
  • P. olsonii: verrucolone
  • P. oxalicum: secalonic acid D & F*, oxaline
  • P. palitans: cyclopiazonic acid*, fumigaclavine A & B*
  • P. paneum: patulin*, roquefortine C*, marcfortines
  • P. polonicum: penicillic acid*, verrucosidin*, nephrotoxic glycopeptides*, cyclopenin, cyclopenol
  • P. roqueforti: roquefortine C*, isofumigaclavine A & B*, PR?toxin*, mycophenolic acid
  • P. solitum: cyclopenin, cyclopenpol. compactins
  • P. verrucosum: ochratoxin A*, citrinin*, verrucolone, verrucins
  • P. viridicatum: xanthomegnin*, viomellein*, vioxanthin*, viridic acid*, penicillic acid*, brevianamide A
  • Eurotium (anamorph Aspergillus subgenus Aspergillus)
  • E. amstelodami: physcion, echinulin
  • E. chevalieri: physcion, echinulin
  • E. herbariorum: physcion, echinulin
  • Petromyces and Neopetromyces (anamorph Aspergillus subgenus Circumdati)
  • Neopetromyces and section Circumdati:
  • A. ochraceus: penicillic acid*, ochratoxin A*, xanthomegnin*, viomellein*, vioxanthin*
  • Petromyces and section Flavi :
  • Petromyces alliaceus: ochratoxin A*, kojic acid
  • A. flavus: kojic acid, 3?nitropropionic acid*, cyclopiazonic acid*, aflatoxin B1*, aspergillic acid
  • A. oryzae: kojic acid, cyclopiazonic acid*, 3?nitropropionic acid*
  • A. nomius: kojic acid, tenuazonic acid*, aflatoxin B1*, B2*, G1*, G2*, aspergillic acid
  • A. parasiticus: kojic acid, aspergillic acid, aflatoxin B1*, B2*, G1*, G2*
  • A. sojae: kojic acid, aspergillic acid
  • Aspergillus Section Nigri:
  • A. niger: naphtho?4?pyrones, malformins, ochratoxin A* (few isolates)
  • A. carbonarius: naphtho-4-pyrones, ochratoxin A*
  • Neosartorya and Hemicarpenteles? (anamorphs in Aspergillus subgenera Fumigati and Clavati)
  • Neosartorya and section Fumigati:
  • N. fischeri: verrucologen*, fumitremorgin A & B*
  • A. fumigatus: gliotoxin*, verrucologen*, fumitremorgin A & B*, fumitoxins**, fumigaclavines* tryptoquivalins
  • Hemicarpenteles and section Clavati:
  • A. clavatus: patulin*, ascladiol, cytochalasin E*, tryptoquivalins
  • Emericella (anamorph Aspergillus subgenus Nidulantes, section Nidulantes, Versicolores and Usti)
  • E. nidulans: sterigmatocystin*, nidulotoxin**, penicillin***
  • A. versicolor: sterigmatocystin*, nidulotoxin**
  • A. ustus: austamide*, austdiol*, austins*, austocystins*
  • Fennellia (anamorphs in Aspergillus subgenus Flavipedes andTerrei)
  • A. terreus: terrein, patulin*, citrinin*, citreoviridin*, territrem*


1. Eurotiales
2. Sphaeriales
3. Dothidiales
4. Unknown

2. Sphaeriales

  • Microascus (anamorph Scopulariopsis)
  • S. brevicaulis: ?
  • Chaetomium
  • C. globosum: chaetoglobosins*, chetomin*
  • Hypocrea (anamorph Trichoderma)
  • T. harzianum: chrysophanol, koninginin A, trichorzianines A + B
  • T. virens: gliotoxin*, gliovirin, viridin
  • T. viride: alamethicins, emodin*, suzukacillin, trichodermin*, trichotoxin A
  • Gibberella (anamorph Fusarium section Arthrosporiella, Discolor, Elegans, Gibbosum, Liseola, Roseum, Sporotrichiella)
  • F. acuminatum: antibiotic Y, chlamydosporol, trichothecenes type A*, enniatins, moniliformin
  • F. avenaceum: antibiotic Y, chlamydosporol, fusarin C, moniliformin, enniatins
  • F. crookwellense: culmorin, fusarin C, trichothecenes type B*, zearalenone*, butenolide, chrysogine
  • F. culmorum: culmorin, fusarin C, trichothecenes type B*, zearalenone*, butenolide, chrysogine
  • F. equiseti: equisetin, fusarochromanone*, trichothecenes type A & B*, zearalenone*, chrysogine
  • F. graminearum: culmorin, fusarin C, trichothecenes type B*, zearalenone*butenolide, chrysogine
  • F. semitectum: equisetin, zearalenone*, chryso-gine, fusapyrone
  • F. oxysporum: fusaric acid, naphthoquinone pigments, nectriafurone, moniliformin, gibepyrones
  • F. poae: fusarin C, trichothecenes type A & B*, butenolide, gamma-lactones,
  • F. proliferatum: fumonisins*, fusaric acid, fusarin C, moniliforme, naphthoquinone pigments, beauvericins, fusaproliferin, fusapyrone
  • F. subglutinans: fusaric acid, fusarin C, moniliforme, naphthoquinone pigments, beauvericin, fumonisin*
  • F. sambucinum: trichothecenes type A*, butenolide, enniatins
  • F. sporotrichioides: fusarin C, trichothecenes type A*, butenolide
  • F. tricinctum: antibiotic Y, fusarin C, butenolide, chlamydosporol, chrysogine, visoltricin
  • F. verticillioides: fumonisins*, fusaric acid, fusarin C, moniliforme, naphthoquinone pigments, gibepyrones
  • Haematonectria (anamorph Fusarium section Martiella)
  • F. solani: fusaric acid, naphthoquinone pigments
  • Claviceps (anamorph Sphacelia)
  • C. purpurea: ergotalkaloids


1. Eurotiales
2. Sphaeriales
3. Dothidiales
4. Unknown

3. Dothidiales

  • Mycosphaerella (anamorph Cladosporium)
  • C. herbarum: epi? and fagi?cladosporic acid**
  • Pleospora (anamorph Stemphylium)
  • S. botryosum: ?
  • Lewia (anamorph Alternaria)
  • A. cf. Alternata: alternariol, alternariol monomethylether, altertoxins
  • A. tenuissima: alternatriol, alternariol monomethylether, altertoxins, tenuazonic acid*
  • Melanopsamma (anamorph Stachybotrys)
  • S. chartarum: satratoxin G and H*


1. >Eurotiales
2. Sphaeriales
3. Dothidiales
1. Eurotiales
2. n connection to known ascomycetes

  • Unknown teleomorph (anamorph Epicoccum)
  • E. nigrum: ?
  • Unknown teleomorph (anamorph Wallemia)
  • W. sebi: Walleminol A & B


1. Eurotiales
2. Sphaeriales
3. Dothidiales
4. Unknown

Notes

* Mycotoxins in the restricted sense used in this chapter;
** not structure elucidated;
*** penicillin is not classified as a mycotoxin, but should not be present in foods.